PATRICIA BACKWELL

PUBLICATIONS

 

Milner RNC, Jennions MD, Backwell PRY (In Press)Keeping up appearances: male fiddler crabs wave faster in a crowd. Biology Letters (accepted Nov 2011)

PDF

Callander C, Backwell PRY, Jennions MD (In Press) Context-dependent male mate choice: the effects of competitor presence and competitor size. Behavioral Ecology (Online)

PDF

Callander C, Jennions MD, Backwell PRY (2012) The effect of claw size and wave rate on female choice in a fiddler crab. Journal of Ethology 30:151-155

PDF

Booksmythe I, Hayes C, Jennions MD, Backwell PRY. 2011) The effects of neighbor familiarity and size on cooperative defense of fiddler crab territories. Behavioral Ecology (Online)

PDF

Callander S, Jennions MD, Backwell PRY. (2011) Female Choice over short and long distances: neighbour effects. Behavioural Ecology & Sociobiology 65:2071-2078

PDF

Bolton J, Callender S, Jennions MD, Backwell PRY. (2011). Even weak males help their neighbours: defence coalitions in a fiddler crab. Ethology (Online)

PDF

Richard N. C. Milner, Michael D. Jennions, and Patricia R. Y. Backwell (2011) Keeping up appearances: male fiddler crabs wave faster in a crowd. Biology Letters (Online)

PDF

Slatyer, RA , Mautz, BS, Backwell PRY, Jennions MD (2011) The genetic benefits of polyandry: a meta-analysis. Biolo. Rev. (Online)

PDF Milner, RNC, MD Jennions, PRY Backwell (2011)  Know thine enemy’s neighbour: neighbour size affects floater’s choice of whom to fight. Behavioural Ecology. 22(5):947-950
PDF

Milner, RNC, MD Jennions, PRY Backwell (2011) Nonindependent mate choice in a fiddler crab: a case of stimulus enhancement. Behavioral Ecology and Sociobiology.65(7):1419-1424

PDF Mautz, Brian; Detto, Tanya; Wong, Bob; Kokko, Hanna; Jennions, Michael; Backwell, Patricia (2011) Male fiddler crabs defend multiple burrows to attract additional females. Behavioural Ecology.22(2):261-267
PDF

Booksmythe I, Jennions MD, Backwell PRY (2011) Male fiddler crabs prefer conspecific females during simultaneous, but not sequential, mate choice. Animal Behaviour 81(4):775-778

PDF Milner RNC, MD Jennions, PRY Backwell (2011). Eavesdropping in crabs: an agency for lady detection. Biology Letters 6(6):755-757
PDF

Booksmythe I, MD Jennions, PRY Backwell 2010. Interspecific assistance: fiddler crabs help heterospecific neighbours in territory defence. Biology Letters 6:748-750

PDF Detto T, Jennions MD, Backwell PRY (2010) When and why do territorial coalitions occur? Experimental evidence from a fiddler crab. Americal Naturalist 175:E119-125
PDF Mokhlesi A, Kamrani E, Backwell PRY, Sajjadi M (2011) Study on the behaviour of two fiddler crabs, Uca sindensis and Uca annulipes (Decapoda: Ocypodidae) in Bandar Abas, Iran. Journal of the Marine Biology Association 91(1):245-249
PDF Milner, RNC, I Booksmythe, MD Jennions, PRY Backwell (2010) Battle of the sexes? Territory acquisition and defence in male and female fiddler crabs. Animal Behaviour 79:735-738
PDF Milner RNC, Detto T, Jennions MD, Backwell PRY (2010) Experimental evidence for a seasonal shift in the strength of a female mating preference. Behavioural Ecology 21:311-316
PDF Milner RNC, Jennions MD, Backwell PRY (2010) Safe sex: male-female coalitions and pre-copulatory mate guarding in a fiddler crab. Biology Letters. 6:180-182
PDF BooksmytheI, Jennions MD, Backwell PRY 2010) Investigating the 'dear enemy' phenomenon in the territorial defence of the fiddler crab Uca mjoebergi. Animal Behaviour. 79:419-423
PDF Booksmythe I, Milner RNC, Jennions MD, Backwell PRY (2010) How do weaponless male fiddler crabs avoid aggression? Behavioural Ecology & Sociobiology. 64:485-491
PDF Milner, RNC, Detto T, Jennions MD, Backwell PRY (2010) Hunting and predation in a fiddler crab. Journal of Ethology. 28:171-173
PDF Murai M, Backwell PRY, Jennions MD (2009) The cost of reliable signalling: experimental evidence for a predictable variation among males in a cost-benefit trade-off between sexually selected traits. Evolution 63(9): 2363-2371
PDF Detto T, Backwell PRY (2009) The fiddler crab Uca mjoebergi uses ultraviolet cues in mate choice but not aggressive interactions. Animal Behaviour. 78: 407-411
PDF Lailvaux SP, Reaney LT, Backwell PRY (2009) Regenerated claws signal performance and fighting ability in the fiddler crab Uca mjoebergi. Functional Ecology 23:359-366
PDF Detto T & Backwell PRY (I2009) Social monogamy in the fiddler crab Uca capricornis. Crustacean Biology 29(3):283-289
PDF Slatyer RA; Fok ESY; Hocking R; Backwell PRY (2008) Why do fiddler crabs build chimneys? Biology Letters.4: 616-618
PDF Booksmythe I; Detto T; Backwell PRY (2008) Fenale fiddler crabs settle for less: cost-benefit tradeoffs in mate choice. Animal Behaviour 76:1775-1781.
PDF

Milner R; Jennions MD; Backwell PRY (2008) Does the environmental context of a signalling male influence his attractiveness? Animal Behaviour. 76:1565-1570

PDF Kinnear M; Smith LMA; Maurer G; Backwell PRY; Linde CC. (In Press). Polymorphic microsatellite loci for paternity analysis in the fiddler crab Uca mjoebergi. Journal of Crustacean Biology. 29(2):273-274
PDF Detto T; Hemmi JM; Backwell PRY. 2008. Colouration and colour changes of the Fiddler Crab Uca capricornis: A Descriptive study. PLoS ONE 3(2):e1629
PDF Fayed SA, Jennions MD & Backwell PRY. 2008. What factors contribute to the Ownership Advantage? Biology Letters. 4:143-145
PDF Reaney L, Sims RA, Sims SWM, Jennions MD & Backwell PRY. 2008. Experiments with robots explain synchronised courtship in a fiddler crab. Current Biology 18:R62-63
PDF Reaney L, Milner R, Detto, T & Backwell PRY. 2007. The effects of claw regeneration on territory ownership and mating success in the fiddler crab Uca mjoebergi. Animal Behaviour. 75:1437-1478
PDF Backwell PRY, Matsumasa M, Double M, Roberts A, Murai M, Keogh JS, Jennions MD. 2007. What are the consequences of being left-clawed in a predominantly right-clawed fiddler crab?. Proceedings of the Royal Society Series B 274:2732-2729 
PDF Reaney L &  Backwell PRY. 2007. Temporal constraints and female preference for burrow width in the fiddler crab, Uca mjoebergi. Behavioral Ecology and Sociobiology 61:1515-1521
PDF Reaney L &  Backwell PRY.2007. Risk-talking behaviour predicts aggression and mating success in a fiddler crab. Behavioral Ecology 18: 521-525
PDF Reading K &  Backwell PRY. 2007. Can beggars be choosers? Male mate choice in a fiddler crab. Animal Behaviour 74:867-872
PDF Murai M & Backwell PRY. 2006. A conspicuous courtship signal in a fiddler crab Uca perplexa: female choice based on display structure. Behavioral Ecology and Sociobiology 60: 736-741
PDF Detto T, Backwell PRY, Hemmi J, Zeil J. 2006. Visually mediated species and neighbour recognition in fiddler crabs (Uca mjoebergi and Uca capricornis). Proceedings of the Royal Society Series B 273:1661-1666
PDF Zeil J, Hemmi J, Backwell PRY. 2006. Fiddler crabs. Current Biology 16: R40-R41
PDF Backwell PRY, Jennions MD, Wada K, Murai M, Christy J. 2006. Synchronous waving in two species of fiddler crabs. Acta Ethologica 9: 22-25
PDF Christy JH, Backwell PRY. 2006.No preference for exaggerated courtship signals in a sensory trap. Animal Behaviour 71: 1239-1246
PDF Murai M & Backwell PRY. 2005. More signalling for earlier mating: conspicuous
male claw waving in the fiddler crab Uca perpelexa.
Animal Behaviour 70: 1093-1097
PDF Morrell LJ, Backwell PRY, Metcalfe NB. 2005. Fighting in fiddler  crabs Uca mjoebergi: what determines duration? Animal Behaviour 70: 653-662
PDF Backwell PRY, Jennions MD. 2004. Coalition among male fiddler crabs. Nature 430: 317 (Reproduced with permission)  
  PDF

Jennions MD, Backwell PRY, Murai M, Christy JH. 2003. Hiding behaviour in  fiddler crabs: How long should prey hide in response to a potential predator? Animal Behaviour 66: 251-257

PDF 

Christy JH, Baum J, Backwell PRY. 2003. Attractiveness of sand hoods built by courting male fiddler crabs Uca musica: test of a sensory trap hypothesis. Animal Behaviour 66: 89-94

PDF  

de Rivera CE, Backwell PRY, Christy JH, Vehrencamp SL. 2003. Density affects female and male searching in the fiddler crab, Uca beebei. Behavioral Ecology and Sociobiology 53: 72-83

PDF

Christy JH, Backwell PRY, Schober U. 2003. Inter-specific attractiveness of structures built by courting male fiddler crabs: experimental evidence of a  sensory trap. Behavioral Ecology and Sociobiology 53: 84-91

PDF

Christy JH, Backwell PRY, Goshima S & Kreuter T. 2002. Sexual selection for structure building by courting male fiddler crabs: an experimental study of behavioral mechanisms. ? Behavioral Ecology 13: 366-374

PDF

Koga T, Backwell PRY, Christy JH, Murai M, Kasuya E. 2001. Male-biased predation of a fiddler crab. Animal Behaviour 62: 201-207

PDF

Christy JH, Backwell PRY, Goshima S. 2001. The design and production of a sexual signal: Hoods and hood building by male fiddler crabs Uca musica. Behaviour 138: 1065-1083

PDF

Backwell PRY, Christy JH, Telford SR, Jennions MD, Passmore NI. 2000. Dishonest signalling in a fiddler crab. Proceedings of the Royal Society Series B 267: 719-724 

PDF

Backwell PRY, Jennions MD, Passmore NI, Christy JH. 1999. Female choice in the synchronously waving fiddler crab, Uca annulipes. Ethology 105: 415-422  (Reproduced with the permission of Blackwell Wissenschafts-Verlag)

 

Christy JH, Goshima S, Backwell PRY, Kreuter TJ. 1998. Nemertean predation on the tropical fiddler crab Uca musica. Hydrobiologia 365: 233-239

PDF

Backwell PRY, O'Hara PD, Christy JH. 1998. Prey availability and selective foraging in shorebirds. Animal Behaviour 55: 1659-1667

PDF

Jennions MD, Backwell PRY. 1998. Variation in courtship rate in the fiddler crab Uca annulipes: is it related to male attractiveness? Behavioral Ecology 9: 605-611  (Reproduced with the permission of Oxford University Press)  

PDF

Koga T, Backwell PRY, Jennions MD, Christy JH. 1998. Elevated predation risk changes mating behaviour and courtship in a fiddler crab Uca beebei Proceedings of the Royal Society Series B 265: 1385-1390  (Reproduced with permission)

PDF

Backwell PRY, Jennions MD. Passmore NI, Christy JH. 1998. Synchronized courtship in fiddler crabs. Nature 391: 31-32  (Reproduced with permission)  

PDF

Jennions MD & Backwell PRY. 1996. Residency and size affect fight duration and outcome in the fiddler crab, Uca annulipes. Biological Journal of the Linnean Society 57: 293-306  (This material has been published in the above journal, the only definitive repository of the content that has been certified and accepted after peer review. Copyright and all rights therein are retained by The Linnean Society of London. This material may not be copied or reposted without explicit permission. For this and other articles go to http://www.idealibrary.com).

PDF

Backwell PRY & Passmore NI. 1996. Time constraints and multiple choice criteria in the sampling behaviour and mate choice of the fiddler crab, Uca annulipes. Behavioral Ecology and Sociobiology 38: 407-416

 

Christy JH & Backwell PRY. 1995. The sensory exploitation hypothesis. Trends in Ecology and Evolution 10: 417 (Reproduced with permission. Copyright owned by Elsevier Science)

PDF

Jennions MD, Bishop PJ, Backwell PRY, Passmore, NI. 1995. Call rate variability and female choice in the African frog, Hyperolius marmoratus. Behaviour 132: 709-720  

PDF

Jennions MD, Backwell PRY, Passmore NI. 1995. Repeatability of mate choice: the effect of size in the African painted reed frog,  Hyperolius marmoratus. Animal Behaviour 49: 181-186

PDF

Backwell PRY, Jennions MD, Christy J & Schobe U. 1995. Pillar-building in the fiddler crab Uca beebei: evidence for a condition-dependent ornament. Behavioral Ecology and Sociobiology 36: 185-192  (The original publication is available on LINK at http://link.springer.de/ Reproduced with permission of Springer-Verlag)

PDF

Polakow D, Backwell PRY, Caithness N, Jennions MD. 1995. Stabilising or directional selection in signalling systems: investigations in a population of painted reed frogs, Hyperolius marmoratus.  South African Journal of Science 91: 270-271  

PDF

Backwell PRY, Jennions MD. 1993. Mate choice in the Neotropical frog, Hyla ebraccata: sexual selection, mate recognition and signal selection. Animal Behaviour 45: 1248-1250

PDF

Jennions MD, Backwell PRY, Passmore NI. 1992. Breeding behaviour of the African frog, Chiromantis xerampelina: multiple spawning and polyandry. Animal Behaviour 44: 1091-1100

PDF

Jennions MD, Backwell PRY. 1992. Chorus size influences on the anti-predator behaviour of a Neotropical frog. Animal Behaviour 44: 990-992 (This material has been published in the above journal, the only definitive repository of the content that has been certified and accepted after peer review.  Copyright and all rights therein are retained by The Association for the Study of Animal Behaviour. This material may not be copied or reposted without explicit permission. For this and other articles go to http://www.idealibrary.com).

 

Backwell PRY & Passmore NI. 1991. Satellite behavior in the leaf-folding frog, Afrixalus delicatus. Journal of Herpetology 25: 497-498 

 

Backwell PRY & Passmore NI. 1991. Sonic complexity and mate localization in the leaf-folding frog, Afrixalus delicatus. Herpetologica 47: 226-229

 

Backwell PRY, Bishop PJ, Passmore NI. 1991. Anuran phonotaxis experiments - does the speaker affect accuracy? Journal of Herpetology 25: 231-232

 

Backwell PRY & Passmore NI. 1991. Advertisement calls and female phonotaxis in Natal dwarf Afrixalus. Journal of African Zoology 4: 275-280

 

Backwell PRY, Passmore NI. 1990. Aggressive interactions and inter-male spacing in choruses of the leaf-folding frog, Afrixalus delicatus. S Afr J Zool 25: 133-137 

 

Backwell PRY & Passmore NI. 1990. Polyandry in the leaf-folding frog, Afrixalus delicates. Herpetologica 46: 7-10 

 

Backwell PRY, Passmore NI. 1990. Suitable approach perches affect female phonotaxis in an arboreal frog. Herpetologica 46: 11-14 

 

Backwell PRY. 1988. Functional partitioning in the two-part call of the Leaf-Folding Frog, Afrixalus brachycnemis. Herpetologica 44: 1-7